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COHO SALMON <br />The life history of Coho Salmon is described in detail in Pacific Salmon Life Histories (Groot <br />and Margolis, 1991) and is included herein by reference. A summary to assist in the <br />discussion of effects of the proposed action is included below. <br />Coho Salmon occur on the Pacific Coast from central California to Alaska. In Puget <br />Sound, Coho generally return to their natal rivers in late summer and autumn after one <br />winter at sea. Coho generally spawn from mid October though early January (Salmon <br />Watcher). After an average incubation period of 42 to 56 days, fry emerge and spend up <br />to 15 months in fresh water prior to migrating to the ocean. Fresh water, Coho depend <br />on visual cues for locating food which primarily of insects. Once leaving rivers and <br />estuaries, Coho enter the Pacific ocean where they range widely for on average a period <br />of sixteen months before returning to spawn and die. <br />STEELHEAD TROUT <br />Steelhead (Oncorhynchus mykiss) occurs on the Pacific Coast from southern California to <br />Alaska (NOAA Species of Concern). The life history of the Steelhead is more complex <br />than any other species of anadromous fish (Barnhart 1986). These fish can be <br />anadromous or fresh water resident. Resident fish are called rainbow trout while the <br />anadromous strain are known as Steelhead (NOAA-NWFSC tech memo-27, NOAA <br />Species of Concern). Steelhead generally spend from one to four years in fresh water prior <br />to smoltification, and up to three years in salt water prior to first spawning. The average <br />spawning age of these fish is four years (NOAA Species of Concern). In addition, <br />Steelhead have the ability to spawn more than once (iteroparity). With the exception of 0. <br />clarki, all other species of Oncorhynchus spawn once and then die (semelparity) (Barnhart <br />1986, NOAA-NWFSC tech memo-27, NOAA Species of Concern). Steelhead may be <br />divided in to two categories, either winter run or summer run fish (Barnhart 1986, <br />California Department of Fish and Game 1954, NOAA-NWFSC tech memo-27, NOAA <br />Species of Concern). Of the two categories, winter Steelhead are the most common and <br />occur in all coastal rivers of Washington, Oregon, and California (NOAA-NWFSC tech <br />memo-27). Winter Steelhead enter freshwater in a sexually mature state between <br />November and April, and spawn soon thereafter in April and May (Barnhart 1986, <br />NOAA Species of Concern). Summer Steelhead are less abundant than winter Steelhead. <br />These fish enter freshwater as immature fish between May and October and do not <br />mature and spawn for several months, generally until January and February (Barnhart <br />1986, NOAA Species of Concern). Steelhead from both winter and summer runs may <br />enter freshwater in the spring or fall and are then called spring or fall -run Steelhead. In <br />some large rivers, Steelhead may enter freshwater throughout the year (Barnhart 1986, <br />California Department of Fish and Game 1954). After emergence from their eggs, <br />Steelhead fry spend from one to four years in freshwater before becoming smolts and <br />migrating to saltwater (Barnhart 1986). <br />